This is the large scale model of turtle visual cortex (the "NGU model")
described in:
Nenadic, Z., Ghosh, B.K. and Ulinski. P. (2003)
Propagating Waves in Visual Cortex: A Large Scale Model of Turtle Visual
Cortex", J. Computational Neuroscience 14:161-184.
and
Nenadic, Z., Ghosh, B.K. and Ulinski. P. (2002) Modeling and Estimation
Problems in the Turtle Visual Cortex, IEEE Trans. Bio-Med. Eng., 49:753-762
It is also described in considerable detail in the file
TurtleVisCortex-descrip.pdf, which is included in this archive.